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Paul Nicklen
Tiny crustaceans called krill, seen here feeding on phytoplankton, form the base of the ocean's food web.
A young man in Oahu, Hawai‘i, gets a tattoo the traditional way, with a sharp comb dipped in ink tapped into the skin. The art of tattoo is practiced across the Polynesian Triangle, with designs and meanings that vary from culture to culture.
Surfers prepare to warm up before a competition on the Hawaiian island of O'ahu. A story in the February 2015 issue documented the importance of surfing in native Hawaiian culture and history.
A manatee swims in Crystal River Reserve, Florida. An April 2013 story documented the tension between humans and manatees, which are threatened by boats, entanglement, and algae blooms.
A polar bear peers into a cabin in Svalbard, the Arctic archipelago in Norway, which was the subject of an April 2009 story. About 3,000 polar bears live in Svalbard, outnumbering the human population.
An Inuit hunter gazes out from the edge of melting sea ice.
Alaska, like many northern latitudes, is warming at around twice the global average. The seas around the state, with rising temperatures, are providing ideal conditions for harmful algal blooms (HABs). These are having a devastating impact on food chains, effectively turning shellfish into poison pills that have claimed the lives of whales, walruses, countless birds and other sea creatures that depend on them. Here, a biologist struggles with her emotions as she calms a sea otter dying a gravel beach in the town of Homer.
Atlantic walruses migrate from Russia to Svalbard, Norway, where they will spend the summer. Svalbard, an archipelago that lies between Norway and the North Pole, has seen dramatic temperature increases and ice melt in the past 50 years due to global warming.
A trio of emperor penguins is surrounded by chicks on the frozen Ross Sea, Antarctica. In a colony of thousands, emperor penguins use unique vocalisations to find and identify their partners and chicks.